1. Hubei Hongshan Laboratory , Wuhan, 430070 , China
2. College of Life Science and Technology, Huazhong Agricultural University , Wuhan, Hubei 430070 , China
3. Shenzhen Institute of Nutrition and Health, Huazhong Agricultural University , Shenzhen 518000 , China
4. Shenzhen Branch, Guangdong Laboratory for Lingnan Modern Agriculture , Shenzhen 518000 , China
5. Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences , Shenzhen 518000 , China