1. Institute of Molecular Physiology and Biotechnology of Plants (IMBIO), University of Bonn , 53115 Bonn , Germany
2. Leibniz Institute of Plant Genetics and Crop Plant Research (IPK) , Department Physiology and Cell Biology, 06466 Seeland, OT Gatersleben , Germany
3. Institute of Crop Science and Resource Conservation (INRES), University of Bonn , 53113 Bonn , Germany