1. Graduate School of Bioagricultural Sciences, Nagoya University, Chikusa, Nagoya, Japan
2. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo, Tokyo, Japan
3. Core Research for Evolutional Science and Technology (CREST), Kawaguchi, Saitama, Japan
4. Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan