1. Transmission, Infection, Diversification & Evolution Group, Max Planck Institute for the Science of Human History, Kahlaische Straße 10, Jena 07745, Germany
2. Centre for Computational Evolution, University of Auckland, 3 Symonds St, Auckland Central, Auckland 1010, New Zealand
3. Department of Linguistic and Cultural Evolution, Max Planck Institute for the Science of Human History, Kahlaische Straße 10, Jena 07745, Germany
4. ARC Centre of Excellence for the Dynamics of Language, ANU College of Asia and the Pacific, The Australian National University, Coombs Building, Liversidge St, Acton ACT 2601, Australia