1. Department of Biosystems Science and Engineering, ETH Zürich, Mattenstrasse 26, 4058 Basel, Switzerland
2. Swiss Institute of Bioinformatics (SIB), Quartier Sorge - Batiment Genopode, 1015 Lausanne, Switzerland
3. Department of Zoology, University of Oxford, Peter Medawar Building South Parks Road Oxford, OX1 3SY, UK
4. Department of Environmental Sciences, ETH Zürich, Universitätsstrasse 16, 8092 Zürich, Switzerland
5. Centre for Computational Evolution, University of Auckland, New Zealand
6. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, 2200 Osborn Dr., Ames, IA 50011 USA