1. Liaoning Key Laboratory for Biological Invasions and Global Changes, College of Bioscience and Biotechnology, Shenyang Agricultural University , Shenyang 110866 , China
2. School of the Environment and Safety Engineering, Jiangsu University , Zhenjiang 212013 , China
3. Department of Biology, University of Konstanz , Konstanz , Germany
4. Zhejiang Provincial Key Laboratory of Plant Evolutionary Ecology and Conservation, Taizhou University , Taizhou 318000 , China