1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences , Mengla 666303 , China
2. Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences , Mengla 666303 , China
3. College of Life Sciences, University of Chinese Academy of Sciences , Beijing 100049 , China
4. College of Agriculture and Forestry, Puer University , Puer 665000 , China
5. College of Bioscience and Biotechnology, Liaoning Key Laboratory for Biological Invasions and Global Changes, Shenyang Agricultural University , Shenyang 110866 , China