1. Division of Metabolism, Endocrinology and Nutrition, Veterans Affairs Puget Sound Health Care System and University of Washington, 1660 South Columbian Way, Seattle, 98108, Washington, USA
2. Oulu Center for Cell-Matrix Research, Biocenter Oulu and Faculty of Biochemistry and Molecular Medicine, University of Oulu, Pentti Kaiteran Katu 1, Linnanmaa, Oulu, Finland