1. RIKEN Center for Sustainable Resource Science , Yokohama, Kanagawa 230-0045 , Japan
2. Institute of Natural Medicine, University of Toyama , Toyama, Toyama 930-0194 , Japan
3. Graduate School of Life and Environmental Science, University of Tsukuba , Tsukuba, Ibaraki 305-8572 , Japan
4. Tsukuba-Plant Innovation Research Center (T-PIRC), University of Tsukuba , Tsukuba, Ibaraki 305-8572 , Japan