1. Graduate School of Agriculture, Kyoto University, Kitashirakawa Oiwake-cho, Sakyo-ku, Kyoto, Japan
2. Department of Forest Molecular Genetics and Biotechnology, Forestry and Forest Products Research Institute, 1 Matsunosato, Tsukuba, Ibaraki, Japan
3. Department of Biology, University of Hawai‘i at Hilo, 200 W. Kāwili St. Hilo, HI, USA