1. Graduate School of Agriculture, Kyoto University , Kitashirakawa Oiwake-cho, Sakyo-ku, Kyoto 606-8502 , Japan
2. Institute of Arctic Climate and Environment Research , Research Institute for Global Change, , Yokohama 236-0001 , Japan
3. Japan Agency for Marine-Earth Science and Technology , Research Institute for Global Change, , Yokohama 236-0001 , Japan