1. Department of Watershed Sciences and Ecology Center, Utah State University, Logan, UT 84322, USA
2. Smithsonian Environmental Research Center, 647 Contees Wharf Road, Edgewater, MD 21037, USA
3. Biology Department, Bryn Mawr College, Bryn Mawr, PA 19010, USA
4. Jackson Estuarine Laboratory, Department of Natural Resources and the Environment, University of New Hampshire, 85 Adams Point Road, Durham, NH 03824, USA