1. Department of Biodiversity, Ecology and Evolution, Faculty of Biology, Universidad Complutense de Madrid , Madrid , Spain
2. Institute of Evolutionary Biology (CSIC—Universitat Pompeu Fabra) , Barcelona , Spain
3. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science , Berlin , Germany
4. Department of Ecology, Evolution, and Organismal Biology, Iowa State University , Ames, IA , United States