1. CSIRO Land & Water, Canberra, ACT 2601, Australia
2. Research School of Biology, Australian National University, Canberra, ACT 2601, Australia
3. School of Biology & Environment Science, University College Dublin, Belfield, Dublin 4, Ireland
4. Earth Institute, University College Dublin, Belfield, Dublin 4, Ireland
5. School of Science and Health & Hawkesbury Institute of the Environment, Western Sydney University, Penrith, NSW 2751, Australia
6. School of Natural Sciences, University of Tasmania, Hobart, TAS 7001, Australia