1. Unité de recherche Œnologie EA 4577, Institut des Sciences de la Vigne et du Vin, University of Bordeaux, USC 1366 INRA, Bordeaux INP, 33140 Villenave d'Ornon, France
2. School of Agriculture, Food and Wine, The University of Adelaide, PMB 1, Glen Osmond, SA 5064, Australia
3. The Australian Wine Research Institute, PO Box 197, Glen Osmond, Adelaide, South Australia 5064, Australia
4. ENSCBP, Bordeaux INP, 33600 Pessac, France
5. Bordeaux Sciences Agro, 33170 Gradignan, France