1. School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, Leicestershire, LE12 5RD, United Kingdom
2. Institute of Biological, Environmental and Rural Sciences, Gogerddan Campus, University of Aberystwyth, Aberystwyth, Ceredigion, SY23 3EB, United Kingdom
3. Department of Chemical Engineering and Biotechnology, University of Cambridge, Phillipa Fawcet Drive, Cambridge, Cambridgeshire, CB3 0AS, United Kingdom