1. Department of Environmental and Biological Sciences, University of Eastern Finland, P.O. Box 1627, 70211, Kuopio, Finland
2. Terrestrial Ecology Section, Department of Biology, University of Copenhagen, Copenhagen Ø 2100, Denmark
3. Center for Permafrost (CENPERM), Department of Geosciences and Natural Resource Management, University of Copenhagen, Copenhagen K 1350, Denmark
4. Ramboll, Niemenkatu 73, 15140, Lahti, Finland
5. Department of Ecological Science, Faculty of Science, Vrije Universiteit Amsterdam, 1081 HV Amsterdam, The Netherlands
6. Natural Resources Institute Finland, Juntintie 154, 77600 Suonenjoki, Finland