1. Department of Biodiversity and Conservation, Real Jardín Botánico de Madrid (RJB-CSIC) , 28014 Madrid , Spain
2. Departament of Plant Biology and Ecology, Faculty of Pharmacy, Universidad de Sevilla , 41012 Sevilla , Spain
3. Island Ecology and Evolution Research Group, Instituto de Productos Naturales y Agrobiología (IPNA-CSIC) , 38206 San Cristóbal de La Laguna , Tenerife, Canary Islands , Spain
4. Naturalis Biodiversity Center , 2333 Leiden , The Netherlands
5. Groningen Institute for Evolutionary Life Sciences, University of Groningen , 9700 AB Groningen , The Netherlands
6. Department of Biology (Botany), Faculty of Sciences, Universidad Autónoma de Madrid , 28049 Madrid , Spain
7. Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid , 28049 Madrid , Spain
8. Department of Biology, Aarhus University , 8000 Aarhus C , Denmark
9. Centre for Functional Ecology, TERRA Associate Laboratory, Department of Life Sciences, University of Coimbra , 3000-456 Coimbra , Portugal