1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences , Kunming 650201, Yunnan , China
2. Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences , Kunming 650201, Yunnan , China
3. University of the Chinese Academy of Sciences , Beijing 100049 , China
4. Institute of Molecular Plant Sciences, School of Biological Sciences, University of Edinburgh , Edinburgh EH9 3JH , UK
5. Royal Botanic Garden Edinburgh , Edinburgh EH3 5LR , UK
6. Lijiang Forest Biodiversity National Observation and Research Station, Kunming Institute of Botany, Chinese Academy of Sciences , Lijiang 674100, Yunnan , China