1. College of Life Sciences, South-Central University for Nationalities, Wuhan, China
2. School of Integrative Plant Science, Cornell University, Ithaca, NY, USA
3. Experiment Center of Forestry in North China, Chinese Academy of Forestry, Beijing, China
4. School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai, China
5. Department of Plants and Crops, unit HortiCell, Faculty of Bioscience Engineering, University of Ghent, Ghent, Belgium