1. Institute of Molecular, Cell and Systems Biology, University of Glasgow, Bower Building, Glasgow G12 8QQ, UK
2. SynthSys - Synthetic and Systems Biology, School of Biological Sciences, University of Edinburgh, Roger Land Building, The King's Buildings, Mayfield Road, Edinburgh EH9 3JD, UK
3. Department of Biology, University of York, Wentworth Way, York YO10 5DD, UK