1. BreMarE - Bremen Marine Ecology , Marine Zoology, , PO Box 330440, 28334 Bremen , Germany
2. Universität Bremen , Marine Zoology, , PO Box 330440, 28334 Bremen , Germany
3. Department of Mathematics and Statistics , Faculty of Science, , 26 Richmond Street, Glasgow, G1 1XH United Kingdom
4. University of Strathclyde , Faculty of Science, , 26 Richmond Street, Glasgow, G1 1XH United Kingdom
5. Oceans & Coasts Research Branch, Department of Forestry, Fisheries, and the Environment , PO Box 52126, Victoria & Alfred Waterfront, Cape Town 8002 , South Africa
6. Bayworld Centre for Research & Education , 5 Riesling Road, Constantia, Cape Town 7806 , South Africa
7. Oceanography Department and Marine Research Institute, University of Cape Town , Private Bag X3, Rondebosch, Cape Town 7701 , South Africa
8. Department of Biological Sciences and Marine Research Institute, University of Cape Town , Private Bag X3, Rondebosch, Cape Town 7701 , South Africa