1. Univ. Lille, CNRS, Univ. Littoral Cote d'Opale, UMR 8187, LOG, Laboratoire d’Océanologie et de Géosciences, 62930 Wimereux, France
2. Institute of Marine Biology, National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 202, Taiwan
3. Equipe BPA, USC ANSES, EA 7394 – ICV – Institut Charles Viollette, Univ. Littoral Côte d'Opale, Quai Robert Masset, 62200 Boulogne sur Mer, France
4. Department of Science and Environment, Roskilde University, DK-4000 Roskilde, Denmark