1. State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, 650223, China
2. Kunming Police Dog Base of the Chinese Ministry of Public Security, Kunming, 650204, China
3. Harbin Police Dog Training Centre, Heilongjiang General Station of Exit and Entry Frontier Inspection, Harbin, 150000, China
4. Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences, Kunming, 650223, China