1. Department of Research, Conservation and Collections, Desert Botanical Garden, Phoenix, AZ, USA
2. Department of Geosciences, University of Arizona, Tucson, AZ, USA
3. Estuary Research Center, Shimane University, Matsue, Shimane, Japan
4. School of Health, Medical and Applied Sciences, Central Queensland University, Townsville, QLD, Australia
5. Department of Botany, University of Wyoming, Laramie, WY, USA