1. RIKEN Center for Sustainable Resource Science , 1-7-22 Suehiro-cho, Tsurumi-ku, Yokohama, 230-0045 Japan
2. Institute of Plant Science and Resources, Okayama University , 2-20-1 Chuo, Kurashiki, 710-0046 Japan
3. School of Life Science and Technology, Tokyo Institute of Technology , 4259 Nagatsuta-cho, Midori-ku, Yokohama, 226-8502 Japan
4. Research Institute for Agriculture and Life Sciences, Tokyo University of Agriculture , 1-1-1 Sakuragaoka, Setagaya-ku, Tokyo, 156-8502 Japan
5. Graduate School of Agriculture and Life Science, The University of Tokyo , 1-1-1 Yayoi, Bunkyo-ku, Tokyo, 113-0032 Japan
6. Institute for Advanced Research, Nagoya University , Furo-cho, Chikusa-ku, Nagoya, 464-8601 Japan