1. Biological Resources Division, Japan International Research Center for Agricultural Sciences, 1-1 Ohwashi, Tsukuba, Ibaraki 305-8686, Japan
2. Japan Science and Technology Corporation, 4-1-8 Motomachi, Kawaguchi, Saitama 332-0012, Japan
3. Laboratory of Plant Molecular Biology, RIKEN Tsukuba Institute, 3-1-1 Koyadai, Tsukuba, Ibaraki 305-0074, Japan
4. Plant Functional Genomics Group, RIKEN Genomic Sciences Center, 3-1-1 Koyadai, Tsukuba, Ibaraki 305-0074, Japan