1. Department of Plant Sciences, University of Oxford , South Parks Rd, Oxford OX1 3RB , UK
2. Department of Biology, Division of Biochemistry, Friedrich–Alexander University Erlangen–Nürnberg , Staudtstrasse 5, Erlangen 91ß58 , Germany
3. BASF Belgium Coordination Center CommV, Innovation Center Gent , Technologiepark-Zwijnaarde 101, Gent 9052 , Belgium
4. Max Planck Institute of Molecular Plant Physiology , Am Mühlenberg 1, Potsdam 14476 , Germany