1. Institute of Agrobiological Sciences, National Agriculture and Food Research Organization, Tsukuba, 305-8604, Japan
2. Institute for Agro-Environmental Sciences, National Agriculture and Food Research Organization, Tsukuba, 305-8604, Japan
3. Ishikawa Prefectural University, Ishikawa 921-8836, Japan
4. Institute of Crop Science, National Agriculture and Food Research Organization, Tsukuba, 305-8518, Japan