1. CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, The Innovative Academy of Seed Design of Chinese Academy of Sciences , Wuhan 430074 , China
2. Hubei Hongshan Laboratory , Wuhan, 430070 , China
3. University of Chinese Academy of Sciences , 19A Yuquanlu, Beijing 100049 , China
4. The New Zealand Institute for Plant & Food Research Ltd, Mt Albert Research Centre , Private Bag, Auckland 92169 , New Zealand
5. School of Biological Sciences, University of Auckland , Private Bag 92019, Auckland 1142 , New Zealand