1. Apple technology innovation center of Shandong Province, College of Horticulture Science and Engineering Shandong Agricultural University Tai‐An 271018 China
2. CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Hubei Hongshan Laboratory The Innovative Academy of Seed Design of Chinese Academy of Sciences Wuhan 430074 China
3. College of Biology and Oceanography Weifang University Weifang 261061 China