1. BESE Division, Plant Cell and Developmental Biology, King Abdullah University of Science and Technology , Thuwal, Kingdom of Saudi Arabia
2. BESE Division, The BioActives Lab, King Abdullah University of Science and Technology , Thuwal, Kingdom of Saudi Arabia
3. Institute of Cytology and Genetics , Novosibirsk 630090, Russian Federation, Russia
4. Novosibirsk State University , Novosibirsk 630090, Russian Federation, Russia
5. Plant Systems Physiology, Radboud University , 6500 AJ Nijmegen, the Netherlands