1. Department of Bioscience, School of Biological and Environmental Sciences, Kwansei Gakuin University , Sanda, Hyogo 669-1330 , Japan
2. Graduate School of Biostudies, Kyoto University , Kyoto 606-8502 , Japan
3. Field Science Center for Northern Biosphere, Muroran Marine Station, Hokkaido University , Muroran 051-0013 , Japan