1. Department of Life Sciences, Pohang University of Science and Technology, Pohang 37673, Korea
2. School of Biological Sciences, Seoul National University, Seoul 151-747, Korea
3. School of Interdisciplinary Bioscience and Bioengineering, Pohang University of Science and Technology, Pohang 37673, Korea
4. Graduate School of Science and Technology, Nara Institute of Science and Technology, Takayama 8916-5, Ikoma, Nara 630-0192, Japan