1. Department of Life Sciences, POSTECH Biotech Center, Pohang University of Science and Technology, Pohang 790-784, Korea
2. School of Interdisciplinary Bioscience and Bioengineering, Pohang University of Science and Technology, Pohang 790-784, Korea
3. RIKEN Center for Sustainable Resource Science, Tsurumi, Yokohama 230-0045, Japan