1. Department of Horticulture, School of Horticulture and Landscape Architecture, Yangzhou University , Yangzhou 225009 , China
2. Joint International Research Laboratory of Agriculture and Agri-Product Safety, the Ministry of Education of China, Yangzhou University , Yangzhou 225009 , China
3. Jiangsu Key Laboratory for Horticultural Crop Genetic Improvement, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences , Nanjing 210014 , China