1. DOE-Plant Research Laboratory, Michigan State University , East Lansing, MI 48824 , USA
2. Donald Danforth Plant Science Center , St. Louis, MO 63132 , USA
3. United States Department of Agriculture, Agriculture Research Service , St. Louis, MO 63132 , USA
4. Department of Biochemistry and Molecular Biology, Michigan State University , East Lansing, MI 48824 , USA
5. Department of Plant Biology, Michigan State University , East Lansing, MI 48824 , USA