1. College of Agriculture and Biotechnology , Zhejiang University, Zijingang Campus, Hangzhou 310058, China
2. Zhejiang University Zhejiang Provincial Key Laboratory of Horticultural Plant Integrative Biology, , Zijingang Campus, Hangzhou 310058, China
3. Zhejiang University State Agriculture Ministry Laboratory of Horticultural Plant Growth, Development and Quality Improvement, , Zijingang Campus, Hangzhou 310058, China
4. School of Biosciences Division of Plant and Crop Sciences, , University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5RD, UK