1. College of Ecology and Environment, Chengdu University of Technology , Chengdu 610059 , China
2. Institute of Wetland Ecology & Clone Ecology/Zhejiang Provincial Key Laboratory of Evolutionary Ecology and Conservation/College of Life Science, Taizhou University , Taizhou 318000 , China
3. Department of Biology, BioCost Group, Faculty of Science, Universidade da Coruña , 15071 A Coruña , Spain