1. Department of Biological Sciences, Louisiana State University, Baton Rouge, Louisiana 70803;
2. The James Hutton Institute, Aberdeen AB15 8QH, Scotland, United Kingdom
3. Department of Plant Pathology, Physiology, and Weed Science, Virginia Polytechnic Institute and State University, Blacksburg, Virginia 24061
4. Division of Biological Sciences, University of Montana, Missoula, Montana 59812
5. The Institute on Ecosystems, University of Montana, Missoula, Montana 59812