1. Seto Marine Biological Laboratory, Field Science Education and Research Center, Kyoto University, Nishimuro, Wakayama, Japan
2. Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa, Chiba, Japan
3. Graduate School of Human and Environmental Studies, Kyoto University, Sakyo, Kyoto, Japan
4. Sesoko Station, Tropical Biosphere Research Center, University of the Ryukyus, Motobu, Okinawa, Japan