1. School of Nature Conservation, Beijing Forestry University, Beijing, China
2. School of Life Sciences, Sun Yat-sen University, Guangzhou, China
3. Ministry of Education Key Laboratory for Biodiversity Sciences and Ecological Engineering, College of Life Sciences, Beijing Normal University, Beijing, China
4. Department of Biological and Environmental Sciences, School of Agricultural, Life and Natural Sciences, Alabama A&M University, Normal, AL, USA
5. Department of Animal and Plant Sciences, University of Sheffield, Sheffield, UK