1. Department of Evolutionary Studies of Biosystems, School of Advanced Sciences, SOKENDAI (The Graduate University for Advanced Studies), Shonan Village, Hayama, Kanagawa 240–0193, Japan
2. Department of Biology, School of Biological Sciences, Tokai University, Minamisawa 5–1–1–1 Minami-ku, Sapporo, Hokkaido 005–8601, Japan
3. Department of international and regional studies, Faculty of Education, Hokkaido University of Education, 1–2 Hachiman, Hakodate, Hokkaido 040–8567, Japan
4. Rikkyo University, 3–34–1 Nishi-Ikebukuro, Toshima-ku, Tokyo 171–8501, Japan