1. CSIRO Agriculture and Food, Queensland Biosciences Precinct, 306 Carmody Rd , St Lucia, Queensland 4067 , Australia
2. The University of Queensland, School of Agriculture and Food Sciences , St Lucia, Queensland 4072 , Australia
3. International Maize and Wheat Improvement Centre (CIMMYT) , Carretera México-Veracruz Km 45, El Batán, Texcoco, México, CP 56237 , Mexico