1. Key Laboratory of Saline-alkali Vegetation Ecology Restoration (Northeast Forestry University), Ministry of Education , Harbin 150040 , China
2. College of Life Sciences, Northeast Forestry University , Harbin 150040 , China
3. Horticultural Sub-academy of Heilongjiang Academy of Agricultural Sciences , Harbin 150040 , China
4. Institute for Sustainable Agroecosystem Services, Graduate School of Agricultural and Life Sciences, The University of Tokyo , Midoricho, Nishitokyo, Tokyo, 188-0002 , Japan