1. Division of Genetics and Genomics, The Roslin Institute, The University of Edinburgh, Midlothian EH25 9RG, UK
2. MRC Integrative Epidemiology Unit, University of Bristol, Bristol BS8 2BN, UK
3. Department of Genetics and Genomic Sciences, Institute of Genomics and Multiscale Biology, Icahn School of Medicine at Mount Sinai, New York, NY 10029, USA
4. Integrated Cardio Metabolic Centre (ICMC), Karolinska Institutet, Huddinge 141 57, Sweden
5. Computational Biology Unit, Department of Informatics, University of Bergen, Bergen N-5020, Norway