1. College of Food Science and Engineering, Tianjin University of Science and Technology , Tianjin 300457 , P. R. China
2. College of Biotechnology, Tianjin University of Science and Technology , Tianjin 300457 , P. R. China
3. Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences , Tianjin 300308 , P. R. China
4. Key Laboratory of Engineering Biology for Low-Carbon Manufacturing, Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences , Tianjin 300308 , P. R. China