1. Huazhong Agricultural University National Key Laboratory for Germplasm Innovation and Utilization of Horticultural Crops, College of Horticulture and Forestry, , Wuhan 430070, China
2. Hubei Hongshan Laboratory , Wuhan 430070, China
3. Institute of Fruit Tree Research, Guangdong Academy of Agricultural Sciences Key Laboratory of South Subtropical Fruit Biology and Genetic Resource, , Guangzhou 510640, China