1. Laboratory of Vaccine Materials, Center for Vaccine and Adjuvant Research, and Laboratory of Gut Environmental System, National Institutes of Biomedical Innovation, Health and Nutrition (NIBIOHN), Osaka, Japan
2. Division of Veterinary Science, Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Osaka, Japan
3. Department of Microbiology and Immunology, Graduate School of Medicine, Kobe University, Hyogo, Japan
4. Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan
5. Graduate School of Medicine and Graduate School of Dentistry, Osaka University, Suita, Osaka, Japan
6. Division of Mucosal Vaccines, International Research and Development Center for Mucosal Vaccines, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan