1. Experimental Evolution Research Group, Max Planck Institute for Evolutionary Biology, Plön, Germany
2. Department of Evolutionary Theory, Max Planck Institute for Evolutionary Biology, Plön, Germany
3. Department of Evolutionary Genetics, Max Planck Institute for Evolutionary Biology, Plön, Germany
4. Department of Veterinary Medicine, Cambridge Veterinary School, University of Cambridge, Cambridge, United Kingdom
5. Department of Genetics, Evolution and Environment, University College London, London, United Kingdom